Portal:Devonian

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The Devonian Portal

Map of the Earth during the late Devonian, c. 370 Ma.

The Devonian (/dɪˈvni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician period.

Fishes, especially jawed fish, reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The armored placoderms began dominating almost every known aquatic environment. In the oceans, cartilaginous fishes such as primitive sharks became more numerous than in the Silurian and Late Ordovician. Tetrapodomorphs, which include the ancestors of all four-limbed vertebrates (i.e. tetrapods), began diverging from freshwater lobe-finned fish as their more robust and muscled pectoral and pelvic fins gradually evolved into forelimbs and hindlimbs, though they were not fully established for life on land until the Late Carboniferous. (Full article...)

Selected Devonian Article

Montage of trilobite genera: Top row: Walliserops, Phacops and Cambropallas; bottom row: Isotelus, Kolihapeltis and Ceratarges

Trilobites (/ˈtrləˌbts, ˈtrɪlə-/; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period (521 million years ago) and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized mineralised exoskeleton, they left an extensive fossil record. The study of their fossils has facilitated important contributions to biostratigraphy, paleontology, evolutionary biology, and plate tectonics. Trilobites are placed within the clade Artiopoda, which includes many organisms that are morphologically similar to trilobites, but are largely unmineralised. The relationship of Artiopoda to other arthropods is uncertain.

Trilobites evolved into many ecological niches; some moved over the seabed as predators, scavengers, or filter feeders, and some swam, feeding on plankton. Some even crawled onto land. Most lifestyles expected of modern marine arthropods are seen in trilobites, with the possible exception of parasitism (where scientific debate continues). Some trilobites (particularly the family Olenidae) are even thought to have evolved a symbiotic relationship with sulfur-eating bacteria from which they derived food. The largest trilobites were more than 70 centimetres (28 in) long and may have weighed as much as 4.5 kilograms (9.9 lb). (Full article...)

Selected Devonian land plant article

Reconstruction of Rhynia gwynne-vaughanii
The rhyniophytes are a group of extinct early vascular plants that are considered to be similar to the genus Rhynia, found in the Early Devonian (around 419 to 393 million years ago). Sources vary in the name and rank used for this group, some treating it as the class Rhyniopsida, others as the subdivision Rhyniophytina or the division Rhyniophyta. The first definition of the group, under the name Rhyniophytina, was by Banks, since when there have been many redefinitions, including by Banks himself. "As a result, the Rhyniophytina have slowly dissolved into a heterogeneous collection of plants ... the group contains only one species on which all authors agree: the type species Rhynia gwynne-vaughanii". When defined very broadly, the group consists of plants with dichotomously branched, naked aerial axes ("stems") with terminal spore-bearing structures (sporangia). The rhyniophytes are considered to be stem group tracheophytes (vascular plants). (Full article...)

Selected Devonian formation

The Tully Formation is a geologic unit in the Appalachian Basin. The Tully was deposited as a carbonate rich mud, in a shallow sea at the end of the Middle Devonian. Outcrops for the Tully are found in New York State and Pennsylvania. It is also found subsurface in western Maryland and northern West Virginia. A number of fossil remains from marine organisms maybe found in Tully out crops. (Full article...)


Model_of_Dunkleosteus_terrelli_(fossil_placoderm)_(Late_Devonian;_Cleveland,_Ohio,_USA)_1_(34189080296)

Selected Devonian fish article

Dipterus valenciennesi

Dipterus (from Greek: δίς dís, 'two' and Greek: πτερόν pteron 'wing') is an extinct genus of lungfish from the middle Devonian period of Europe and North America. In most respects, Dipterus, which was about 35 centimetres (14 in) long, closely resembled modern lungfish. Like its ancestor Dipnorhynchus, it had tooth-like plates on its palate instead of real teeth. However, unlike its modern relatives, in which the dorsal, caudal, and anal fin are fused into one, Dipterus's fins were still separated.

The genus was established by Adam Sedgwick & Roderick Murchison in the year 1828. (Full article...)

Selected Devonian invertebrate

Solitary rugose coral Grewingkia canadensis in three views; Ordovician, Indiana

The rugosa, also called the tetracorallia or horn coral, are an extinct order of solitary and colonial corals that were abundant in Middle Ordovician to Late Permian seas.

Solitary rugosans (e.g., Caninia, Lophophyllidium, Neozaphrentis, Streptelasma) are often referred to as horn corals because of a unique horn-shaped chamber with a wrinkled, or rugose, wall. Some solitary rugosans reached nearly a meter in length. However, some species of rugose corals could form large colonies (e.g., Lithostrotion). When radiating septa were present, they were usually in multiples of four, hence Tetracorallia in contrast to modern Hexacorallia, colonial polyps generally with sixfold symmetry.

Rugose corals have a skeleton made of calcite that is often fossilized. Like modern corals (Scleractinia), rugose corals were invariably benthic, living on the sea floor or in a reef-framework. Some symbiotic rugose corals were endobionts of Stromatoporoidea, especially in the Silurian period. Although there is no direct proof, it is inferred that these Palaeozoic corals possessed stinging cells to capture prey. They also had tentacles to help them catch prey. Technically they were carnivores, but prey-size was so small they are often referred to as microcarnivores. (Full article...)

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